Pei, Heng Pei, Lim Kuang Hock, Cheong Yoon Ling, Noraini Osman, Noraina Majid, Tan Poai Ean, Shyamini Ann Samson, et al. , trans. “The Collections of the Two Native, Near Threatened Pteropid Species in Peninsular Malaysia by the Institute for Medical Research, Malaysia: The Large Flying Fox (Pteropus Vampyrus) and Island Flying Fox (Pteropus Hypomelanus)”. Journal of Wildlife and Parks 36 (September 25, 2021): 59–74. Accessed September 16, 2026. https://jwp.wildlife.gov.my/index.php/jwp/article/view/62.